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Desire asked Many plants have stomata that take in CO2 at night and release during the day. Why is this form of photosynthe?
an advantage for plants living in a hot, dry climate?
And got the following answer:
Plants have to compromise between water loss in hot climates with intense sunlight and freely exchanging gases because they do not have separate pathways for water movement and gas exchange with the atmosphere. Both water and CO2 are rate limiting factors for photosynthesis to proceed and both pass through the stomata. To get enough CO2 in C3 photosynthesis too much water may be lost to transpiration and the plant injured by desiccation. The plant must shut off the stomata to prevent desiccation but then it cannot photosynthesize for the lack of CO2. By adding the separating step C4 plants pull the CO2 in and move it quickly away so the CO2 is always flowing in faster than in C3 plants. This is what requires the extra energy, moving the CO2 faster so more can diffuse down the gradient the plant creates. The CO2 cannot be lost once internalized so this allows CAM plants to separate the CO2 acquisition in time as well as space. Cam plants regulate their stomatal opening times to be nocturnal so the can gather and store the CO2 to allow photosynthesis to proceed with daylight. This makes C4 transpiration ~25% of a C3 plant, so they conserve water. The extra cost of C4 does the plant no good in lower light & cooler, moister conditions but allows plants to survive in the tropical grasslands. Some plants switch from C3 to CAM depending on habitat factors. CAM Bromeliads are rainforest epiphytes where water can be erratically available in the tree canopy. After a rainy period the plant has water and operates with the energy efficient C3. When the plant runs out of water it switches to CAM and opens the stomata at night to sequester the CO2 with less water loss. Then with daylight the plant can photosynthesize again.